Since 2014

Guest scientist at the Laboratory of Functional and Environmental Ecology-EcoLab Toulouse, France (CNRS, UPS, INPT)

June 2017 - October 2018

Confirmed researcher at the UFZ Departments Conservation Biology (K. Henle)

June 2015-2017

Marie-Curie Fellow at the UFZ Departments Systems ecotoxicology (M. Liess) and Conservation Biology (K. Henle)

November 2012 - December 2013

Researcher at the CNRS Unit EcoLab (UMR 5245), Toulouse (JL Probst)

September 2013

Guest professor in Conservation Biology at the Department of Bioenvironmental Systems Engineering, National Taiwan University

Dec 2007-October 2012

Post-Doc at the biological station for experimental ecology of the CNRS (USR 2936)

July 2007 - Sept 2007

Visiting researcher at the Helmholtz Centre for Environmental Research - UFZ, Department Conservation Biology, Leipzig, Germany

March 2007 - Dec 2007

Post-Doc at the Emirates Center of Wildlife Propagation, Missour, Morocco

Sept 2006 - Feb 2007

Post-Doc at the Helmholtz Centre for Environmental Research - UFZ, Department Conservation Biology, Leipzig, Germany

Sept 2004 - Aug 2006

Temporary Teaching and Assistant Researcher (ATER) in behavioural ecology, Laboratoire d’Ecologie Animale, Université d’Angers, France

  • Research on sexual selection in the Palmate newt, bet-hedging in Rana sp.

  • Teaching ranges from 1st to 4th year students

  • Created a course in Ethology for 1st year students in Psychology

Apr - June 2004

Scientific guide. Parc Zooloqique de Clères, Muséum National
d’Histoire Naturelle, France

Sept 03 - Jan 04

Temporary teacher in high school (biology classes), Region Paris, France

Mar 2002 - Jul 2003

Biological Assistant at the Centre National de la Recherche Scientifique, Laboratoire de Parasitologie Evolutive, Paris, France

  • Microsatellites markers screening by cross-amplification in Starling and Blue tit.

Apr - June 2001

Scientific guide. Parc Zooloqique de Clères, Muséum National d’Histoire Naturelle, France



2001 - 2005

Ph. D. with honors in Behavioural Ecology at the Muséum National
d’Histoire Naturelle, Paris, France
Dissertation title : Sexual selection and honest signalling in the
Common peafowl (Pavo cristatus)
Supervised by Dr. G. Sorci and Dr. M. Saint Jalme
Thesis defence on the 22th June 2005

2000 - 2001

Master’s degree (Grade: B) in Ethology at the Université Paris 13, France

1999 - 2000

Bachelor’s degree (Grade: B)  in Ecosystem and Population Biology at the
Université Pierre et Marie Curie, Paris, France

June 1995

Baccalauréat in Sciences, equivalent to the A level (Grade: A), France



Awards and fellowships

Jun 15- May 17

Marie Skłodowska-Curie – Individual Fellowship, program H2020 (172 k€).
PI of the EU project “FreeMi - Using Freshwater organisms to mitigate the pathogen Batrachochytrium dendrobatidis

Jan 2007

Post-doctoral Fellowship from the RENECO private compagny (100 k€).
PI of the project “Maternal investment in an endangered bird, the houbara bustard”

Apr 04

Award of the best talk, by the ASAB (Association for the Study of Animal Behaviour) ASAB Eastern Meeting, Brighton (UK)

Mar 2004

Traveling Fellowship from the Comité National des Sciences Biologiques, Paris

Mar 2004

Traveling Fellowship from the GDR 2155 Ecologie Comportementale,
Université de Bourgogne




Institutional responsibilities

Since 2017

Expert member of the CSRPN Occitanie (Scientific Regional Council for Natural Heritage), France

Expert delegate for amphibians
Referee expert for the Natural Reverse of Néouvielle (Pyrenees, France)
More than one expertise per month


Administrative responsible and manager of the amphibian facility Station of Experimental Ecology of the CNRS, Moulis (France)
Agreement for animal experiments and national certificate for amphibian housing and breeding

 Reviewing and editing activities

Expert reviewer of funding applications

National Geographic Society
CIFRE doctoral fellowship (France)
Belgium Academy of Science
United States – Israel Binational Science Foundation
Czech Science Foundation (Czech Republic)


Journal editing

Since 2014, editor for Biodiversity and Conservation
Since 2012, editor for Nature Conservation


Article reviewing

I regularly review articles for international scientific journals, including: Ecology Letters, Conservation Letters, Proceedings of the Royal Society London B, Biodiversity and Conservation, BMC Ecology, Animal Behaviour, Behavioral Ecology, Behavioral Ecology and Sociobiology, Plos One, Science of the Total Environment, Journal of Avian Biology, Ethology, Behavioural Processes, Behaviour, Journal of Wildlife Diseases, Biodiversity Data Journal, Revue Suisse de Zoologie, Amphibia-Reptilia, Journal of Visualized Experiments, Herpetological Conservation and Biology, etc.


Book refereeing

I was the scientific referee for the book of M. Gokhale Joshi, Bosses of the wild: Lessons from the corporate jungle, McGraw Hill Education Ed., 2013


Teaching activities

Sep 2013

Guest Professor in Conservation Biology (Chair of Prof. Yu-Pin Lin). National Taiwan University, Taipei, Taiwan

2008 - 2011 

Teaching in Conservation Biology (co-responsible for this module) in 2nd year of Master (Master 2)
Master “Biodiversity, Ecology and Evolution”
University Paul Sabatier, Toulouse, France

Sep 04 - Sep 06 

Temporary Teaching and Research Assistant (ATER) at the University of Angers (France)
100 hours of teaching per year
I was responsible for 5 modules
Students in Ecology (Faculty of Science) and students in Psychology (Faculty of Letters)
Levels: graduate and undergraduate (from 1st year of university to 2nd year of Master)

All types of teaching: lectures, courses, and excursions
Main teaching: Ethology, Animal and Human Behaviour, Behavioural Ecology, Zoology, Ecology, Conservation Biology, Evolutionary Biogeography, Phylogeny, Taxonomy, Comparative Anatomy

2001 and 2004   

Lectures in Conservation Biology and Evolutionary Ecology at the University of Rouen, France
Bachelor students

Sep 03 - Jan 04

Temporary teacher in high school (Maître Axilliaire), biology classes, part time, 94 hours, France
Pupils from 11 to 18 years old

2001 and 2004

Scientific guide at the National Museum of Natural History (MNHN) (>40 hours)
All age classes (from kindergarden to adults)
Zoological Park of Clères, MNHN, Clères, France




  1. The RACE project: Risk Assessment of Chytridiomycosis to European Amphibian Biodiversity
    Conférences Jacques Monod on Emerging Fungal Diseases, Jun 2011, Roscoff, France

  2. Diseases as stressors: amphibian biodiversity threatened by emerging chytridiomycosis
    Stress & Evolution Meeting, Jul 2010, Münster, Germany

  3. Manipulation of maternal investment in a vulnerable species
    Le Réveil du Dodo III, Mar 2009, Montpellier, France

  4. Structural colouration of the eyespots correlates with male mating success in the peacock (Pavo cristatus)
    International Society for Behavioural Ecology, Jul 2006, Tours, France

  5. Maternal differential allocation in peafowl Pavo cristatus
    European Society for Evolutionary Biology, Aug 2005, Krakow, Poland

  6. Multiple sexual advertisements honestly reflect health status in peacocks (Pavo cristatus)
    International Society for Behavioural Ecology, Jul 2004, Jyväskylä, Finland


  1. Head or Tail’ – Impact of biotic environment on functional traits of amphibian larvae (invited)
    Conference Functional Ecology and Environment, Jul 2017, Toulouse, France

  2. Chytridiomycosis in the Pyrenees: general review, mitigation
    34th GEEFSM (Group for the study of ecopathology of mountain wildlife), Oct 2016, Laruns, France

  3. Zooplankton consumers protect amphibians against Bd in Pyrenean lakes
    27th International Congress for Conservation Biology-ECCB, Aug 2015, Montpellier, France

  4. Zooplankton predators protect amphibians against Bd in Pyrenean mountain lakes
    Amphibian Conservation Research Symposium, Apr 2015, Cambridge, UK

  5. The role of zooplankton in the dynamics of infection by Batrachochytrium dendrobatidis
    34th DPG (German Society for Protozoology), Mar 2015, Magdeburg, Germany

  6. Female fruit flies use sophisticated public information to avoid copulating with sperm limited males (invited)
    European Conference on Behavioural Biology, Jul 2012, Essen, Germany

  7. Manipulation of maternal investment in the Houbara bustard, a vulnerable species
    International Society for Behavioural Ecology, Aug 2008, Ithaca, NY, USA

  8. Male sexual attractiveness affects the allocation of maternal resources into the eggs in peafowl
    International Society for Behavioural Ecology, Jul 2006, Tours, France

  9. Multiple sexual advertisements honestly reflect health status in peacocks (Pavo cristatus) (awarded)
    Association for the Study of Animal Behaviour Easter Meeting, Apr 2004, Brighton, UK

  10. Communication sexuelle, signaux multiples et condition dépendance chez le Paon bleu
    SFECA (French Society for the Study of Animal Behaviour), Apr 2003, Tours, France

Distribution of research results to the general public

I consider communication with the general public as full part of my research work. I regularly interact with the general public when I am in the field and also during cultural and scientific events.


Flyers created for the general public

“What we do against chytridiomycosis for the amphibians of the Pyrenees and elsewhere” (French, English and German)


Articles written for the general public

Loyau A & Schmeller DS. 2010. Heiße Parade im Wüstensand. Spektrum der Wissenschaft, Oct, pp18-20
Loyau A. 2004. Le Paon, une espèce fondamentale au Parc Zoologique de Clères. La lettre de la SECAS 39:6

Courtois E., Loyau A. , Schmeller, D.S. (2011) : Les NACs, précautions à prendre pour limiter les risques de propagations des maladies émergentes« exotiques. Semaine Veterinaire 1471: 46


Participation to cultural and scientific events

Oct 2015 

 Conference on light pollution for The day of the Night, Ariège, France

Nov 2010

Interview and meeting with the public at the Research Festival La Novela, Toulouse, France

Jul 2010

Collaboration with a novelist, Hélène Duffau, to write a short story Vous avez dit Euprocte? Published in the books Douze chercheurs en quête d’auteur (Loubatières Ed., 2011)

Jun 2010 

Interview for a documentary produced by BiodivERsA to promote research

Jan and Nov 2010

Public seminar at the Public Meeting of the Human-Environment Observatory of Haut Vicdessos, Auzat, France

2009 and 2010

I had posters the Festival of Science (La Fête de la Science), Foix, France




Participation to the education of school teachers to promote conservation biology and research

May 2015                   Mission « Teachers in the lab », teaching and discussions with around 20 school teachers, EcoLab, Toulouse, France


Interventions in schools to promote conservation biology and research to children

Jan and May 2011    Junior high school Stendhal (Collège Stendhal), Toulouse, France
Jun 2012                     Primary school of Auzat, France


Communication through medias and press coverage

Jul 2013, Interview for a documentary on amphibians and chytridiomycosis for the channel ARTE
Mai 2010, Interview during more than 1 hour for a radio program Radio Mon Païs, Toulouse, France

I also disseminate my research results to the public thanks to the media. Results of my research on chytridiomycosis and animal behaviour (peacocks, drosophila, and houbara bustard) have received considerable attention by newspapers (Die Zeit, Leipziger Volkszeitung , Sächsische Zeitung, Esslinger Zeitung), magazines (e.g. National Geographic, Scientific American, The New Scientist, Discovery magazine, Science et Vie Junior, Spektrum der Wissenschaft), scientific blogs (e.g. Science daily, Science News, Eurek Alert, New Scientist.com, BBC), as well as books written for the public (e.g. F. Flam, The score: How the Quest for Sex Has Shaped the Modern Man, Avery Ed., 2008; P. Ackermann, Masculins singuliers: enquête sur la nouvelle identité des hommes, Laffont Ed., 2009; B. Natterson Horowitz & K. Bowers, Zoobiquity: What Animals Can Teach Us About Being Human, Ebury Ed., 2012; R. Pookottil, Beem: Biological Emergence-Based Evolutionary, Fossil Fish Ed., 2013). I also contribute regular to the blog of P³ to reach out to the general public and students.