My Google Scholar Profile can be found here:

Articles in peer-reviewed journals

  1. Sentenac, H., Loyau, A. , Zoccarato, L., Jassey, V.E.J., Grossart, H.-P., Schmeller, D.S., (2023). Biofilm community composition is changing in remote mountain lakes with a relative increase in potentially toxigenic algae. Water Research, 120547. pdf

  2. Bates, K.A., Friesen, J., Loyau, A., Butler, H., Vredenburg, V.T., Laufer, J., Chatzinotas, A., Schmeller, D.S. , (2023). Environmental and Anthropogenic Factors Shape the Skin Bacterial Communities of a Semi-Arid Amphibian Species. Microbial Ecology 86, 1393–1404. pdf

  3. Schmeller, D.S., Cheng, T., Shelton, J., Lin, C.-F., Chan-Alvarado, A., Bernardo-Cravo, A., Zoccarato, L., Ding, T.-S., Lin, Y.-P., Swei, A., Fisher, M.C., Vredenburg, V.T., Loyau, A. , (2022). Environment is associated with chytrid infection and skin microbiome richness on an amphibian rich island (Taiwan). Scientific Reports 12, 16456. pdf

  4. Schmeller, D.S. , Urbach, D., Bates, K., Catalan, J., Cogălniceanu, D., Fisher, M.C., Friesen, J., Füreder, L., Gaube, V., Haver, M., Jacobsen, D., Le Roux, G., Lin, Y.-P., Loyau, A. , Machate, O., Mayer, A., Palomo, I., Plutzar, C., Sentenac, H., Sommaruga, R., Tiberti, R., Ripple, W.J., (2022). Scientists' warning of threats to mountains. Science of the Total Environment, 158611. pdf;

  5. Mayer, A., Egger, C., Loyau, A. , Plutzar, C., Schmeller, D.S. , Gaube, V. (2022). Mountain pastures increase the resilience of livestock farming to extreme events in the Ariège department, France. Agronomy for Sustainable Development 42: 49.pdf

  6. Bates, K.A., Sommer, U., Hopkins, K.P., Shelton, J.M.G., Wierzbicki, C., Sergeant, C., Tapley, B., Michaels, C.J., Schmeller, D.S. , Loyau, A., Bosch, J., Viant, M.R., Harrison, X.A., Garner, T.W.J., Fisher, M.C. (2022). Microbiome function predicts amphibian chytridiomycosis disease dynamics. Microbiome 10: 44. pdf

  7. Machate, O., Schmeller, D.S., Loyau, A., Paschke, A., Krauss, M., Carmona, E., Schulze, T., Moyer, A., Lutz, K., Brack, W. (2022). Complex chemical cocktail, containing insecticides diazinon and permethrin, drives acute toxicity to crustaceans in mountain lakes. Science of the Total Environment: 154456. pdf

  8. Sentenac H., Loyau A. , Leflaive J., Schmeller D.S. (2022). The significance of biofilms to human, animal, plant and ecosystem health. Functional Ecology 33: 294 - 313. pdf

  9. Haver M., Le Roux G., Friesen J., Loyau A. , Vredenburg V.T., Schmeller D.S. (2021) The role of abiotic variables in an emerging global amphibian fungal disease in mountains. Science of the Total Environment 152735. pdf

  10. Bernardo-Cravo, A., Schmeller, D.S., Chatzinotas, A., Vredenburg, V.T., Loyau, A. , 2020. Environmental Factors and Host Microbiomes Shape Host-Pathogen Dynamics. Trends in Parasitology 36, 29-36. pdf

  11. Thomas, V., Wang, Y., Rooij, P.V., Verbrugghe, E., Baláž, V., Bosch, J., Cunningham, A.A., Fisher, M.C., Garner, T.W.J., Gilbert, M.J., Grasselli, E., Kinet, T., Laudelout, A., Lötters, S., Loyau, A., Miaud, C., Salvidio, S., Schmeller, D.S. , Schmidt, B.R., Sluijs, A.S.-v.d., Steinfartz, S., Veith, M., Vences, M., Wagner, N., Canessa, S., Martel, A., Pasmans, F., (2019). Mitigating Batrachochytrium salamandrivorans in Europe. Amphibia-Reptilia 40 (3), 265-290. pdf

  12. Bridgewater, P., Loyau, A., Schmeller, D.S., (2019). The seventh plenary of the intergovernmental platform for biodiversity and ecosystem services (IPBES-7): a global assessment and a reshaping of IPBES. Biodiversity and Conservation 28 (10): 2457–2461 . DOI pdf

  13. Beukema, W., Martel, A., Nguyen, T.T., Goka, K., Schmeller, D.S. , Yuan, Z., Laking, A.E., Nguyen, T.Q., Lin, C.-F., Shelton, J., Loyau, A. & Pasmans, F. (2018) Environmental context and differences between native and invasive observed niches of Batrachochytrium salamandrivorans affect invasion risk assessments in the Western Palaearctic. Diversity and Distributions 24(12): 1788-1801pdf

  14. Valbuena-Ureña, E., Oromi, N., Soler-Membrives, A., Carranza, S., Amat, F., Camarasa, S., Denoël, M., Guillaume, O., Sanuy, D., Loyau, A., Schmeller, D.S. & Steinfartz, S. (2018) Jailed in the mountains: Genetic diversity and structure of an endemic newt species across the Pyrenees. PLOS ONE, 13, e0200214.pdf

  15. Fisher M.C., Ghosh P., Shelton J.M.G., Bates K., Brookes L., Wierzbicki C., Rosa G.M., Farrer R.A., Aanensen D.M., Alvarado-Rybak M., Bataille A., Berger L., Böll S., Bosch J., Clare F.C., A. Courtois E., Crottini A., Cunningham A.A., Doherty-Bone T.M., Gebresenbet F., Gower D.J., Höglund J., James T.Y., Jenkinson T.S., Kosch T.A., Lambertini C., Laurila A., Lin C.-F., Loyau A. , Martel A., Meurling S., Miaud C., Minting P., Ndriantsoa S., O’Hanlon S.J., Pasmans F., Rakotonanahary T., Rabemananjara F.C.E., Ribeiro L.P., Schmeller D.S., Schmidt B.R., Skerratt L., Smith F., Soto-Azat C., Tessa G., Toledo L.F., Valenzuela-Sánchez A., Verster R., Vörös J., Waldman B., Webb R.J., Weldon C., Wombwell E., Zamudio K.R., Longcore J.E., Garner T.W.J. (2018) Development and worldwide use of non-lethal, and minimal population-level impact, protocols for the isolation of amphibian chytrid fungi. Scientific Reports 8: 7772.pdf

  16. O’Hanlon S.J., Rieux A., Farrer R.A., Rosa G.M., Waldman B., Bataille A., Kosch T.A., Murray K.A., Brankovics B., Fumagalli M., Martin M.D., Wales N., Alvarado-Rybak M., Bates K.A., Berger L., Böll S., Brookes L., Clare F., Courtois E.A., Cunningham A.A., Doherty-Bone T.M., Ghosh P., Gower D.J., Hintz W.E., Höglund J., Jenkinson T.S., Lin C.-F., Laurila A., Loyau A., Martel A., Meurling S., Miaud C., Minting P., Pasmans F., Schmeller D.S., Schmidt B.R., Shelton J.M.G., Skerratt L.F., Smith F., Soto-Azat C., Spagnoletti M., Tessa G., Toledo L.F., Valenzuela-Sánchez A., Verster R., Vörös J., Webb R.J., Wierzbicki C., Wombwell E., Zamudio K.R., Aanensen D.M., James T.Y., Gilbert M.T.P., Weldon C., Bosch J., Balloux F., Garner T.W.J., Fisher M.C. (2018) Recent Asian origin of chytrid fungi causing global amphibian declines. Science 360: 621-627. pdf

  17. Schmeller, D.S., Loyau, A., Bao, K., Brack, W., Chatzinotas, A., De Vleeschouwer, F., Friesen, J., Gandois, L., Hansson, S.V., Haver, M., Le Roux, G., Shen, J., Teisserenc, R., Vredenburg, V.T., (2018). People, pollution and pathogens – global change impacts in mountain freshwater ecosystems. Science of the Total Environment 622–623: 756-763.pdf

  18. Ripple, W.J., Wolf, C., Newsome, T.M., Galetti, M., Alamgir, M., Crist, E., Mahmoud, M.I., Laurance, W.F., Auliya, M., Droste, N., Egli, L., Everaars, J., Ferreira, C., Friese, K., Henle, K., Kühn, I., Loyau, A., Pe'er, G., Priess, J.A., Schiller, J., Schmeller, D.S., Schröter, M., Spangenberg, J.H., Wiemers, M., and more, (2017): World scientists’ warning to humanity: a second notice. Bioscience 67(12): 1026-1028. pdf

  19. Frenken T., Alacid E., Berger S.A., Bourne E.C., Gerphagnon M., Grossart H.-P., Gsell A.S., Ibelings B.W., Kagami M., Küppe F.C., Letche P.M., Loyau A., Miki T., Nejstgaard J.C., Rasconi S., Reñé A., Rohrlack T., Rojas-Jimenez K., Schmeller D.S., Scholz B., Seto K., Sime-Ngando T., Sukenik A., Van de Waal D.B., Van den Wyngaert S., Van Donk E., Wolinska J., Wurzbacher C., Agha R. (2017). Integrating chytrid fungal parasites into plankton ecology. Research gaps and needs. Environmental Microbiology 19(10): 3802–3822. pdf

  20. Schmeller, D.S., Weatherdon, L.V., Loyau, A., Bondeau, A., Brotons, L., Brummitt, N., Geijzendorffer, I.R., Haase, P., Kuemmerlen, M., Martin, C.S., Mihoub, J. B., Rocchini, D., Saarenmaa, H., Stoll, S., Regan, E.C. (in press). A suite of essential biodiversity variables for detecting critical biodiversity change. Biological Reviews 93 (1), 55 - 71. pdf

  21. Courtois, E.A., Loyau, A. , Bourgoin, M., Schmeller, D.S., (2017). Initiation of Batrachochytrium dendrobatidis infection in the absence of physical contact with infected hosts – a field study in a high altitude lake. Oikos 126: 843–851. pdf

  22. Loyau, A. , Schmeller, D.S., (2017). Positive sentiment and knowledge increases tolerance towards conservation actions. Biodiversity and Conservation 26 (2), 461-478. pdf

  23. Loyau, A. , Cornuau, J.H., Clare, F.C., Schmeller, D.S. (2016). Side effects of itraconazole on post-metamorphic Alytes obstetricans after a cold stress. Amphibia Reptilia 37 (2016): 345-357.pdf

  24. Clare, F.C., Daniel, O., Halder, J.B., Bielby, J., Semenov, M.A., Jombart, T., Loyau, A., Schmeller, D.S., Cunningham, A.A., Rowcliffe, M.J., Garner, T.W.J., Bosch, J., Fisher, M.C. (2016). Climate forcing of an emerging pathogenic fungus across a montane multi-host community. Philosophical Transactions of the Royal Society B: Biological Sciences 371 (1709), 20150454. pdf

  25. Dagaeff, A.-C., Pocheville, A., Nöbel, S., Loyau, A. , Isabel, G. & Danchin, É. (2016). Drosophila mate copying correlates with atmospheric pressure in a speed learning situation. Animal Behaviour 121, 163-174. pdf

  26. Germain, M., Blanchet, S., Loyau, A. & Danchin, É. (2016). Mate-choice copying in Drosophila melanogaster: Impact of demonstration conditions and male-male competition. Behavioural Processes 125: 76-84. pdf

  27. Cornuau, J. H., Schmeller, D. S. , Courtois, E. A., Jolly, T., & Loyau, A. (2015). It takes two to tango: relative influence of male and female identity and morphology on complex courtship display in a newt species. Ethology 121 (3): 218–226. pdf

  28. Cornuau, J. H., Schmeller, D. S. , Pigeault, R., Sibeaux, A., Tourat, A., & Loyau, A. (2014). Information-content of morphological and behavioural sexual traits in the Palmate newt (Lissotriton helveticus). Behavioural Processes 108: 36-42. pdf

  29. Schmeller, D. S., Blooi, M., Martel, A., Garner, T. W. J., Fisher, M. C., Azemar, F., Clare, F. C., Leclerc, C., Jäger, L., Guevara-Nieto, M., Loyau, A., & Pasman, F. (2014). Microscopic aquatic predators strongly affect infection dynamics of a globally emerged pathogen. Current Biology, 24 (2): 176-180. pdf
    Equal contributions by the two first and two last authors.

  30. Cornuau, J.H., Schmeller, D.S., Pigeault, R., and Loyau, A. (2014). Resistance of morphological and behavioural sexual traits of the Palmate newt (Lissotriton helveticus) to bacterial lipopolysaccharide treatment. Amphibia Reptilia 35: 63-71.

  31. Courtois, E.A., Cornuau, J.H., Loyau, A., and Schmeller, D.S. (2013). Distribution of Amphibiocystidium sp. in palmate newts (Lissotriton helveticus) in Ariège, France. Herpetology Notes 6: 539-543 pdf

  32. Drechsler A., Geller D., Freund K., Schmeller D.S. , Künzel S., Rupp O., Loyau A., Denoël M., Valbuena-Ureña E., Steinfartz S. (2013): What remains of a 454 run: estimation of success rate of microsatellite loci development in selected newt species (Calotriton asper, Lissotriton helveticus, Triturus cristatus), Ecology and Evolution 3(11): 3947–3957. pdf

  33. Courtois E.A., Lelong C., Calvez O., Loyau A. , Schmeller D.S. (2013): The use of Visible Implant Alpha tags for Anuran tadpoles, Herpetological Review 44(2):231-233. pdf

  34. Loyau A. , Cornuau J.H., Clobert J., Danchin E. (2012): Incestuous sisters: mate preference for brothers over unrelated males in Drosophila melanogaster. PLoS ONE 7(12): e51293. doi:10.1371/journal.pone.0051293pdf

  35. Loyau A. , Blanchet S., Van Laere P., Clobert J., Danchin E. (2012): When not to copy: female fruit flies use sophisticated public information to avoid mated males. Scientific Reports 2:768. pdf DOI: 10.1038/srep00768

  36. Loyau A. , Schmeller D.S. (2012): Mixed reproductive strategies of the Common Moorhen on a micro-scale as revealed by genetic data. Comptes Rendus Biologies: 335 (10-11):673-679. pdf

  37. Schmeller D.S., Henle K., Loyau A. , Besnard A., Henry P-Y. (2012): Bird-monitoring in Europe – a first overview of practices, motivations and aims. Nature Conservation 2:41-57.pdf

  38. Cornuau J., Rat M., Schmeller D.S., Loyau A. (2012): Multiple signaling in the Palmate newt: longer is not better if you do not court, Behavioural Ecology and Sociobiology 66 (7): 1045-1055 pdf

  39. Henle, K., S. Bell, L. Brotons, J. Clobert, D. Evans, C. Goerg, M. Grodzinska-Jurcak, B. Gruber, Y. Haila, P.-Y. Henry, A. Huth, R. Julliard, P. Keil, M. Kleyer, J. Kotze, W. Kunin, S. Lengyel, Y.-P. Lin, A. Loyau , G. Luck, W. Magnuson, C. Margules, Y. Matsinos, P. May, I. Sousa-Pinto, H. Possingham, S. Potts, I. Ring, J. Pryke, M. Samways, D. Saunders, D. Schmeller, J. Simila, S. Sommer, I. Steffan-Dewenter, P. Stoev, M. Sykes, B. Tóthmérész, R. Yam, J. Tzanopoulos, and L. Penev. (2012). Nature Conservation - a new dimension in Open Access publishing bridging science and application. Nature Conservation 1:1-10. pdf

  40. Schmeller D.S. , Loyau A. , Dejean T., Miaud C. (2011): Using amphibians in lab studies – precautions against the emerging infectious disease Chytridiomycosis. Laboratory Animals 45:25-30. DOI Iridescence

  41. Loyau A., Lacroix F. (2010): Watching sexy displays improves hatching success and offspring growth through maternal allocation. Proceedings of the Royal Society of London, Biological Series 277 (1699): 3453-3460. DOI-link. Iridescence

  42. Loyau A., Schmeller D.S. (2009): Polymorphic microsatellites identified by cross-species amplifications in the European Coot Fulica atra. Journal of Ornithology 150 (3): 703 - 707. DOI-link Iridescence

  43. Loyau A. , M. Petrie, M. Saint Jalme, G. Sorci. (2008) : Do peahens not prefer peacocks with more elaborate trains? Animal Behaviour 76 (5):e5-e9.Iridescence

  44. Loyau A. , D. Gomez, B. Moureau, M. Théry, N. S. Hart, M. Saint Jalme, A.T.D. Bennett & G. Sorci. (2007) : Iridescent structurally based coloration of eyespots correlates with mating success in the peacock. Behavioral Ecology 18 (6): 1123 - 1131, DOI-LinkIridescence

  45. Loyau A. , M. Saint Jalme, R. Mauget & G. Sorci. (2007) : Male sexual attractiveness affects investment of maternal resources into the eggs in peafowl (Pavo cristatus). Behavioral Ecology and Sociobiology 61 (7): 1043-1052. DOI-Link

  46. Loyau A., M. Saint Jalme & G. Sorci. (2007) : Non-defendable resources affect peafowl lek organization: a male removal experiment. Behavioural Processes, 74:64-70.lek

  47. Loyau A. & J. Szczepaniak. (2006) : A darting protocol in a large carinate, the peacock (Pavo cristatus). Journal of Wildlife Management, 70 (6):1813-1815. pdf 6
  48. Loyau A. , B. Moureau, M. Richard, P. Christie , P. Heeb and G. Sorci. (2005) : Cross-amplification of polymorphic microsatellites reveals extra-pair paternity and brood parasitism in Sturnus vulgaris. Molecular Ecology Notes, 5:135-139.

  49. Loyau A., M. Saint Jalme & G. Sorci. (2005) : Intra and intersexual selection for multiple traits in the peacock (Pavo cristatus). Ethology, 111:810-820.

  50. Loyau A., M. Saint Jalme, C. Cagniant & G. Sorci. (2005) : Multiple sexual advertisements honestly reflect health status in peacocks (Pavo cristatus). Behavioral Ecology and Sociobiology, 58:552-557.

Other articles

Non-peer reviewed articles

Blooi, M., Schmeller D. S. , Martel A. Garner T. J. W., Fisher M. C. , Clare F. , Loyau A. , and Pasmans F. 2013. Environmental predation of Batrachochytrium dendrobatidis determines infection dynamics. Pages 57-57 in 17th European congress of Herpetology.

Courtois E., Loyau A. , Schmeller, D.S. (2011) : Les NACs, précautions à prendre pour limiter les risques de propagations des maladies émergentes« exotiques », Semaine Veterinaire 1471: 46.pdf
Loyau A. (2004) . Le Paon, une espèce fondamentale au Parc Zoologique de Clères. La lettre de la SECAS, 39:6.Secas

Press coverage

France 3: 18:30 - Les tribulations d'une scientique en montagne

France Inter - Terre au Carre

France3-Report on Bd-work in the Pyrenees

Science et vie Junior Dec. 2012

Osundefender, Dec. 2012

BBC (the original story by Katharine Sanderson), Dec. 2012

Report on the amphibian disease chytridiomycosis in Faszination Wissen of the Bayrisches Fernsehen Sept. 2013 (watch it here).


Disease Pyramid

20minutes, LaDepeche, Le Petit Journal, Medical News Today, Phys.Org, Health Write Up,



Science Perspective, The Scientist, National Geographic, Die Welt, Badische Zeitung, n-TV, CBC, BBC, The Hindu, The Atlantic, New Scientist, Cosmos, The New York Times, Independent, Nature, Washington Post, Natural History Museum, Scinexx, Axios, Spektrum der Wissenschaft, Uppsala Nya Tidning, Free Malaysia Today, Nature World News, Daily Enterpriser, Canadian Homesteading, Immortal News, General Anzeiger, Tagesspiegel, Tiroler Tageszeitung, ZDFheute, Deutschlandradio, Forskning, StandNews, Sveriges Radio,


Micropredators article in Current Biology

Red Orbit, Science Codex, Reforma, Money, IdW, Die Zeit, NewsMedical, EurekAlert, The Scientists, Altwire, MyScience, Blog lajme flash, Colorado News, NewsAmphibiaAustralia, Phys.Org, Science NewsLine, SciGuru, AbstractMagazine, Media NewsWire, Finsights, Broadcast India, PressZoom, ScienceDaily, Torch-Blog, Bremer Nachrichten,, Scientific American, Spektrum der Wissenschaft, Berliner Zeitung (print, internet), Grist,



La Novela

Audio Inteview

OHM-Haut Vicdessos (Podcast - France Culture), site web


Behavior of Drosophila melanogaster

Real Clear Journal Club


2010: Houbara - Proceedings Royal Society London

Science ORF, Discovery magazine, Sify news, DNA India, 7th Space, NetIndia, Oman Observer, Bright Surf, Yahoo - India, Andrha News, India Interacts, Leipziger Internet Zeitung, Scinexx, Deutsche Welle, News Views, Partnership for European Environmental Research (PEER), Africa Geographic, Science daily, Red Orbit, Anne for Carversville, ThaIndian, FirstScience News, Physorg.Com, North Africa Environment News, Tiergesundheit aktuell (Germany) , National Geographic (Ker Than, USA), gooNews (Japan), CalcuttaTube, AllVoices, NewKerala, The Escapist, Epoch Times Deutschland, National Geographic (Italy), LazyBuzz News, Free Daily News, Research in Germany, CNRS, SCIAM (China) ,

Bayrischer Rundfunk - IQ and RadioWissen (MP3)

Leipziger Volkszeitung, Tuesday 29.06.2010

Sächsische Zeitung, 01.07.2010

Esslinger Zeitung, 10.07.2010

Giessener Allgemeine, 10.07.2010

Oberhessischer Bote, 10.07.2010


Loyau A. Behavioural ecologists elucidated how peahens choose their mates, and why.
Blackwell Publishing (2005-08-01)
Eurek Alert (2005-08-01)
Science Daily (2005-08-17)

Scienzz, Scienceticker, Science 2.0, Cosmos Magazine, CNRS,


Kunz K. 2007. Attraktiv = gesund?, Ulmer online

Natércia F. 2007 . Balanço entre força e beleza na cauda do pavão. Cienciae Cultura. CC